Abstract is: Lycaenidae is the second-largest family of butterflies (behind Nymphalidae, brush-footed butterflies), with over 6,000 species worldwide, whose members are also called gossamer-winged butterflies. They constitute about 30% of the known butterfly species. The family comprises seven subfamilies, including the blues (Polyommatinae), the coppers (Lycaeninae), the hairstreaks (Theclinae), and the harvesters (Miletinae).